About Me

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Canberra-based naturalist, conservationist, educator since 1980. I’m passionate about the natural world (especially the southern hemisphere), and trying to understand it and to share such understandings. To that aim I’ve written several books (most recently 'Birds in Their Habitats' and 'Australian Bird Names; origins and meanings'), run tours all over Australia, and for the last decade to South America, done a lot of ABC radio work, chaired a government environmental advisory committee and taught many adult education classes – and of course presented this blog, since 2012. I am the recipient of the Australian Natural History Medallion, the Australian Plants Award and most recently a Medal of the Order of Australia for ‘services to conservation and the environment’. I live happily in suburban Duffy with my partner Louise surrounded by a dense native garden and lots of birds.

Thursday 23 November 2023

Dhilba Guuranda–Innes National Park; a South Australian coastal glory

It took decades, but this spring I finally got back to this park which was a favourite of mine while I lived in Adelaide. Lou had never seen it and, as a coastal aficionado, was instantly hooked!

Dawn at Shell Beach, Dhilba Guuranda–Innes National Park
I spent time there in the 1970s (!), some of it while working part-time at the then Torrens Teachers College (now part of the University of South Australia). It had been declared Innes National Park shortly before that, in 1970, primarily to protect populations of endangered Malleefowl and Western Whipbirds. The change to the current name came fifty years later in 2020, when joint management with the Narungga traditional owners began, albeit somewhat belatedly.

It is at the toe of the foot of Yorke Peninsula, the middle leg-like one of the three peninsulas that dominate the South Australian coastline.

Dhilba Guuranda–Innes National Park is at the end of the red arrow, though even after you
click on it to enlarge it, you'll find it obscured by the label for Adelaide! The next map,
via Google Earth, might be more useful.

Only the dark green on the 'toe' at the end of the arrow is the park. It only covers
9000 hectares of land, but its significance is also made clear on this map, as it is
the only significant area of uncleared land on Yorke Peninsula. The rest has been turned
to wheat production and grazing land. It is worth noting that the larger
Eyre Peninsula to the west has quite a bit more original vegetation,
and a concomitant number of reserves. Map from Google Earth.
This map of the park is taken from the 2023 management plan, very recently adopted. (As is the preference now, it is a fairly slight document compared with its 2003 predecessor, setting out priorities and aims rather than prescriptions.)
The colours refer to the various management zones, which you can look at in the plan
via the link above, but are not very relevant to this post. It might be useful for locating
sites referred to in the following photos, otherwise just ignore it! For the record we
camped at Shell Beach, in the north of the park, the most remote campground we
could find. We would recommend it too.
The current sea level was only reached some 7000 years ago, but immediately prior to that it was 120 metres lower than it currently is. The exposed sands and banks of sea shells were blown by the wind into a blanketing sheet of low dunes, which percolating water transformed over time into aeolianite (ie wind-deposited) limestone. There are still dunes, some loose but most anchored by vegetation, and the underlying limestone can be seen in the cliffs that line the coast, interspersed by sandy beaches.
Ethel Beach (named for a ship that was wrecked there in 1904),
seen from the top of the cliffs.
Looking south from Pondalowie Bay to West Cape with its lighthouse.
An associated feature is the presence of the islets just offshore.
Looking north from West Cape to Pondalowie Bay beyond the headland in the middle of the photo.
Just off the headland is South Islet, and above it is Middle Islet and North Islet (I have to say that
the naming of the coastal features has been less than imaginative, though Pondalowie is the
shining exception).
North Islet (also known as Royston Islet) from Royston Head. This is at the
end of a lovely little walk through the heathland.
Thirty kilometres to the west - about halfway to the southern tip of Eyre Peninsula at the mouth of Spencer Gulf - is Wedge Island, which was farmed from the mid-nineteenth to the mid-twentieth century. It is still in private ownership, but has an important conservation function as a predator-free haven. In particular it supports a translocated population of Critically Endangered Brush-tailed Bettongs (known as Woylies in Western Australia) Bettongia penicillata, from which a population has been recently reestablished in Dhilba Guuranda–Innes, along with Woylies from Western Australia.
Wedge Island, 30km away, from Pondalowie Bay. (On the Google Earth
map above it lies at the end of the upper 'prong' of the red arrow.)
Another result of the earlier inundation of the lowlands by the sea is the series of saline lakes behind the coastal dunes, clearly visible in the park map above. Four of these these lakes supported gypsum mining from 1889 to 1973, and the small town of Inneston from 1927. When the park was declared in 1973 the mined lakes were excluded for possible resumption of mining, and this was still the case by the time of the 2003 management plan. That has since apparently been rectified, but I can't find just when this occurred. 
Lake Brown; it is not named on the map above, but it is almost adjacent to the
Shell Beach campground.

The lakes have locally unique environments, including these samphire flats.
They are fringed with Swamp Paperbarks Melaleuca halmaturorum which,
like the samphire, are highly salt-tolerant.

Stabilised sand dunes are characteristic of the area behind the beaches.

Vegetated dune behind our campsite.
Above the cliffs however are open heathlands, rich in flowering plants.
Heathland splashed yellow with flowering Grounsel Daisy Senecio sp.
at Pondalowie Bay.
Middle Islet in Pondalowie Bay, with flowering Coastal Beard-Heath Leucopogon parviflorus
in the foreground. This large shrub is an important component of the coastal heaths.
Coastal Beard-Heath dominating the walk through the dunes from Shell Beach
campground to the beach.

The interior of the park, which is not readily accessible, is dominated by nine species of mallee eucalypt, some of which also come close to the sea.

Flowering mallee near Shell Beach.

Red Mallee Eucalyptus oleosa by the campground.

At which point I might as well segue into introducing some other flowering plants that we enjoyed, nearly all from different families, starting with a closeup of the Coastal Beard-Heath we met earlier.

Most other beard-heaths that I know are small shrubs, so these huge bushes are most impressive.
    Here are some more conspicuous large shrubs from the heathlands and mallee of the park.
Sea Box Alyxia buxifolia Family Apocyanacae (a large, mostly tropical family with many
highly toxic species). Sea Box is another large shrub found widely in coastal southern Australia.
I love the odd little 'catherine wheel' flowers.
Common Fringe-myrtle Calytrix tetragona. This lovely shrub is found widely in southern
Australia (including our garden), but I never tire of it. Nor apparently do the
numerous insects which are attracted to the flowers.

Coastal Velvet-bush Lasiopetalum discolor Family Malvaceae (in its new vast sense; Lasiopetalum
was formerly in Sterculiaceae). This attractive shrub, common in the park, has an interesting
distribution along the south coast of Western and South Australia, plus north-western Tasmania.

Cocky's Tongue Templetonia retusa, always a favourite of mine with these large glowing
pea flowers, grows in heaths and mallee from south-western WA to as far east in
South Australia as about here  and up to the Flinders Ranges.

The remaining plants I want to share with you are small shrubs or ground covers, and a climber. (What anyone who knows me would also expect are orchids - but sadly, none were to be found. We were there in the third week of September which should have been good, but at the Visitors' Centre we were told that they'd all flowered early and were done! I was looking forward to trying out the excellent-looking new field guide to South Australian orchids, but unfortunately not this time...)

Pigface Carpobrotus rossii, a familiar and tough beach dune sprawler found right along the
Australian south coast.

Sticky Goodenia Goodenia varia; its leathery toothed leaves can be found in coastal
sands and inland mallee across southern Australia.

Coastal Fanflower Scaevola angustata, also in the Family Goodeniaceae, like the goodenia
above. (I had this down as S. linearis, but that is apparently not in the park.)
This species is endemic to South Australia.

Coastal Cushion Bush Leucophyta brownii. This daisy isn't in flower here (though the
Senecio behind it is), but it's a species I remember fondly from my days in this
part of the world. It is so tough, growing out of cracks in the limestone with
the salt spray blowing over it.

Old Man's Beard Clematis microphylla, a familiar, common and widespread climber
in much of Australia, but not to be ignored because of that!
Of the non-feathered animals, the most obvious were the ubiquitous Western Grey Kangaroos Macropus fuliginosus.
 
They are more brown than grey, despite the name. They are found right across southern
Australia from the coast to the semi-arid inland, westward from central NSW and Queensland.
In the east of their range they are seemingly being displaced by the rapidly spreading
Eastern Grey Kangaroos M. robustus.
It was too cold and windy much of the time for reptiles to be active, though the big skinks called Shinglebacks Tiliqua rugosa were starting to move around. We are very fond of these, and I confess that as a child I kept some as pets (in those days it was legal, with no permit required). At one stage in the park I even rescued one that had settled into the shade of someone's camper van, leaving them bemused as they wanted to be moving on.
They are common and unmistakable with their short tail and diamond-shaped head.
They are long-lived and form lifelong pair bonds, most unusually for a reptile.
Which leaves us with the birds, which were many but often a bit on the skulking side. Here are a few to enjoy. The first couple were just by the main road through the park.
This Brush Bronzewing Phaps elegans was the first bird we saw in the park, to my delight.
Not only is it an extremely attractive pigeon, but in my experience it is shy and hard to
approach. It is found both in dense coastal heaths and more inland forests and mallee across
southern Australia, and is always a pleasure to meet.
Emus Dromaius novaehollandiae can be encountered throughout the park, and
clutches of chicks were being tended by their fathers, necessitating even more
than usual care while driving the road. There are actually two clutches here,
and the other somewhat indifferent father (or so I assume) can be seen in the background.
I haven't knowingly seen 'shared parenting' among emus like this before and
a quick search doesn't give me any further information on this behaviour.
The next four species were regulars around the campground.
Fan-tailed Cuckoos Cacomantis flabelliformis call incessantly in spring,
and things were no different in Dhilba Guuranda–Innes!
Grey Currawongs Strepera versicolor in eastern Australia are clearly ashy-grey.
In the mallee lands (from western Victoria to about Adelaide) they are black,
making identification difficult where they overlap with Pied Currawongs.
West of Adelaide this Yorke and Eyre Peninsula subspecies is grey-brown,
and are often called Brown Currawongs.
Male Purple-backed Fairywren Malurus assimilis, a truly glorious bird.
It was long recognised as a separate species, then more recently lumped
with the Variegated Fairywren, but recent more detailed analysis
has returned the situation to where it was from the 19th century.
The Purple-backed is found across a huge area of the inland, while
Variegated is limited to the east coast. The blue-purple on the crown,
which continues onto the back, can be seen here.
Spiny-cheeked Honeyeaters Acanthagenys rufogularis are found across most of
the inland of Australia, plus the south and west coasts. They are a big honeyeater
with a distinctive and evocative fluting call.

There are three species of fieldwren, all in Australia, generally shy birds of low dense vegetation. Of these the Rufous Fieldwren Calamanthus campestris has by far the biggest range, from western Victoria to the middle western coast of Australia, and from the coast to the inland deserts. This was the first time however that I'd managed to lay camera on one, on the walking track to the West Cape lighthouse.

Rufous Fieldwren sitting up against the sun, checking on the intruders.
This one, with the sun behind me, shows the rufous nicely.
And of course there are seabirds and shorebirds; here are three that we enjoyed, all at Shell Beach, our 'local' while we were there, while we sat quietly either by the rocks at the end of the beach, or on the dunes above it.

It's always a particular pleasure to see Hooded Dotterels (or Plovers) Thinornis cucullatus,
not least because they are a species at risk, especially on the east coast, but also listed
as Vulnerable (ie to extinction) both nationally and here in South Australia.
This was was working Shell Beach early in the morning before visitors arrived.
Sooty Oystercatchers Haematopus fuliginosus are also in trouble on the east coast, but
not so much elsewhere. They are supposed to favour rocky platforms, leaving the beaches
to Pied Oystercatchers, but it's surprising how often birds don't read the books!

Pied Cormorants Phalacrocorax varius aren't at all threatened, but I just like this picture.
Of the four inland Australian cormorants, this is the one most likely to be seen in the ocean.
I hope you now feel that this is a park worth visiting (and on the way there the small Yorke Peninsula towns nearly all have excellent bakeries!); just as further encouragement here are a couple more photos of the sublime beaches.
Browns Beach at the northern end of the park.
Pondalowie Bay again, the southern end.

I hope to see you again in three weeks, when I might have something a bit different. Thanks for visiting.

NEXT POSTING THURSDAY 14 DECEMBER
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Thursday 2 November 2023

Costa Rica; where the Americas meet #3. Animals, the tinies and the hairies.

This is the third of a four-part series on Costa Rica. As I explained last time, the planned post to incorporate all the animals except birds got too long, so here is the rest of that one, with the perhaps unlikely pairing of invertebrates (mostly insects) and mammals. If you missed the first post in the series, you may find it helpful with regard to the places mentioned in this post.

Invertebrates often miss out entirely and, while I won't be guilty of that, there's going to be less information here than for the other animal groups, simply because I don't have it, or because it doesn't even seem to exist anywhere accessible. Many I don't even have a name for but hopefully we can enjoy them without that. Here's a small sample of the many 'little animals' that we saw. 

An orbweb spider Argiope submaronica, at Tapirus Lodge. Although it's not very large, one
has been recorded at La Selva Research Station as capturing, wrapping and feeding on,
a Proboscis Bat (see below for some live and healthy ones).

Grasshoppers and katydids are everywhere; here are a couple of unidentified ones we admired. 

In the rain at Tapirus Lodge.
A large striking species that we saw on both visits, this one was in a large group
close to the sea at Tortuguera.

Beetles are a major part of any ecosystem, and here are three ranging from quite small to extremely large!

A beautifully iridescent little chrysomelid (or leaf beetle) on a telescope at the Bougainvillea Hotel.
Perhaps it was trying to look bigger...
Elephant Beetle Megasoma elephas, some 10cm long. Only the males have the extraordinary
'tusk', used for duelling with other males. This beauty and the next had been temporarily relocated
to the garden of the wildlife artist 'Cope' (see the previous post).
Hercules Beetle Dynastes hercules, an equally striking animal, one of the
rhinoceros beetles. A male can be up to 17cm long, though this one was a little less dramatic.
Another remarkable aspect of this beetle is that it can actually fly, one of the largest insects
to do so. Again only the males have the horn. The huge larvae play an important role
in breaking down fallen logs in the forest.
Leafcutter ants are ubiquitous and fascinating (and hard to meaningfully photograph I've found).
This is part of a column, carrying leaves they've cut to the nest where they are used to
cultivate a fungus that the ants eat. Sometimes you can see ants hitching a ride on
the leaves - these are said to be guarding against parasitic wasps.

Part of the above-ground part of a leafcutter ant colony which may cover tens of square
metres which the ants keep clear of vegetation.

Tapirus Lodge maintains a large light trap - a large double-sided screen with lights to attract insects during the night. It's not clear if it's part of a study, or just to showcase the insects, many of which remain on the screen the next day; while there we daily examined this screen carefully, and found some great rewards. This cicada, and a couple of the moths below, are on this background.

We didn't see this lovely cicada anywhere else, but they were probably high above our heads.

A couple of other invertebrates were quite unexpected.

This amazing tailless whip-scorpion (Family Phrynidae) was in someone's room at Tortuguera.
She (the human occupant of the room) was admirably calm about it, and one of our number
was equally admirable in moving it to a tree trunk outside.They are arachnids but not scorpions
(or even true whip scorpions!). They have no sting or venom, but the jaws can puncture skin if
you do something silly (see next photo). They are nocturnal, hiding under bark or living in caves,
found throughout the world's tropics. This one was at least ten centimetres across.
They use the back six legs for walking, and the front two stretch forward as sensory organs.
Closeup of the scary jaws (or pedipalps).
This land crab was encountered while walking on a forest track in the rain at Monteverde,
not what we expected a few hundred metres above sea level and far from the sea.
Unfortunately I can't find out anything about it.

And I know that at least some of you have been waiting somewhat impatiently for the butterflies. Your wait is over. There were of course many, some of them even identifiable, in some cases with some help (thanks Steve!).

Banded Peacock Anartia fatima males duelling - or least the top one was, the bottom one
was just trying to mind his own business. The other was performing aerobatics to
intimidate - while it's not obvious, in the photo he was in the midst of a full reverse
loop in the air! I'm sure there is information on this behaviour available, but
I can't find it. (And it's possible the male was trying to impress a female,
but again I can't find any information to support this.)
 And for the record, here's what the same Banded Peacock looks like from above.

White Peacock Anartia jatrophae in the same garden at Turtle Lodge in Tortuguera
as the Banded Peacocks above.
Isabella's Longwing Eueides isabella, Cerro Lodge near the Pacific central coast.
It is found throughout Central America, tropical South America and the Caribbean.
Crimson-patched Longwing Heliconius erato, Caño Negro, far north.
Owl Butterfly Caligo sp., La Fortuna, one of a genus of large slow butterflies
found throughout the Neotropics. The 'eyes' are supposed to resemble those of owls.

The remaining four, all moths, were resting on or near the Tapirus Lodge light traps; I can only put a name on the first of them.

Orizaba Silkmoth Rothschildia orizaba. This magnificent moth is huge, up to 18cm
across. The four transparent 'window panes' in the wing give rise to its Spanish name, Cuatro Ventanas.

The remaining three must remain anonymous, unless you can help me. They were chosen from many I could have used from the photos I took at the light trap.

Peacock Moth Automeris io, which is found across much of North America.
(Thanks Fanny!)

And, with something of a leap, that brings us to some mammals. Costa Rica has a rich mammal fauna - as with all animal groups there it seems - but of course most are nocturnal and many are hard to find, especially with a group. What follows however is a fairly good array of species, especially for a very small country.

Sloths, along with armadillos and anteaters, are the only surviving old South Americans, whose ancestors long predated the coming of the invaders from the north which now dominate in South America. You can read more on this in the first post in this series, but it means that they get top billing here! Moreover Costa Rica sometimes seems awash with sloths - they are a major tourist attraction and there are some unsavoury stories of tourist facilities moving sloths to their property to attract customers. I have read too that there is said to be a greater biomass of three-toed sloths in Costa Rica than of any other (non-human) mammal. There are two species of sloth in Costa Rica, each belonging to an entirely different family - in fact their last common ancestor (a ground-dwelling animal) lived some 28 million years ago, so they are not closely related, despite looking very similar. Their lifestyle has led to an impressive example of parallel evolution.

Brown-throated Three-toed Sloth Bradypus variegatus, at Tapirus Lodge. And they do seem
to spend a lot of time scratching! All sloths have three toes on their hind feet, and this family
is named for its front feet (look at its 'hand' on the branch). Famously they come down
to the foot of the tree to defecate about once a week - and I've still not read a
plausible explanation for this. However a recent publication, which is also
unimpressed by all other explanations, suggests it is merely that this is what
their ground-living ancestors did, and there's not been enough selection pressure
(ie from predation during the process) to change the habit.. Hmm.
Hoffman's Two-toed Sloth Choloepus hoffmanni, La Selva Research Station.
They are supposed to be fairly common but this is the only one we saw,
and at night. Not a great photo but I chose it because it shows the
two front claws quite clearly. It also has no tail, while the three-toeds
have a short one.
And in a somewhat bizarre aside, the mighty Zoonomia Project, coordinated by university laboratories around the world, recently compared the complete genomes (an extraordinary concept) of 240 mammal species around the world. One aspect was comparing the number of olfactory receptor genes, a pretty good indicator of the sensitivity of a mammal's ability to smell. Not many people, I suspect, would have bet on Hoffman's Two-toed Sloth to come in third in this 'competition' (behind only the African Elephant and the Nine-banded Armadillo). Another Costa Rican came in fourth - read on!

Tapirs evolved in North America, but became extinct there after spreading to South America and Asia. There are three species in the Americas, but only one in Central America. This is Baird's Tapir Tapirus bairdii, which is found throughout Central America and just enters South America in north-western Colombia. It is the biggest of the American tapirs and is found from mangroves to cloud forests. At Tapirus Lodge - appropriately - a family has become used to visiting the lodge for kitchen scraps, though they are totally wild animals and come and go unpredictably.
Baird's Tapir on a walking track at Tapirus Lodge. The typical tapir proboscis,
a soft, flexible snout for grasping food and smelling the air, shows well here.
This is a big animal, weighing up to 300kg.
Everyone wants to see monkeys in the Neotropics of course and Costa Rica has plenty, though only four species. Three of these are fairly easy to see, but the Red-backed Squirrel Monkey occurs only in two national parks on the Pacific Coast, and unfortunately we didn't see them.
Central American (or Geoffroy's) Spider Monkey Ateles geoffroyi in the lodge garden
at Caño Negro in the far north, though we saw them in several places. Though spindly
with seemingly impossibly long slender limbs and tail, it is regarded as one of the
largest American monkeys, weighing up to 9kg (though this is a young one). Small
groups forage for fruit during the day, and come together for the night roost.
This unusual photo of a mother and baby Mantled Howler Monkey Alouatta palliata was
made possible by the series of suspension bridges above deep rainforest gullies on the
walking tracks at Selvatura Park at Monteverde. This big monkey is found throughout
Central America and down the north-west coast of South America to Ecuador, living
in quite large groups and living primarily on leaves. These are a very low energy
food source, though of course there are plenty of them! To me the rushing roar of
the male howler monkey is one of the sounds of the tropical American forests,
and it can be heard kilometres away.
White-faced Capuchin Cebus capucinus, at Cerro Lodge on the middle Pacific Coast, though
it occurs throughout the country. A fairly small monkey, it travels in groups and eats almost
anything, from fruit and flowers to insects and small lizards. Local guides (especially bird guides)
are likely to describe them as 'bad monkeys' as they actively hunt bird nests, and many breeding
attempts fail due to them.
Microbats (ie the small insect-hunters, not the big fruit bats) are rarely easy to see but two Costa Rican species often roost during the day in conspicuous sites, in groups of one male and a 'harem' of females, and in very distinctive postures. Both are less than 5cm long.
Greater White-lined Bats Saccopteryx bilineata in an alcove behind the fridge in
the open-walled restaurant at Esquinas Lodge in the far south rainforests. Their
'normal' site would be a hollow tree or between tree buttresses. This is their
characteristic pose, upside down with raised head and propped on their forearms.

Proboscis Bats Rhynchonycteris naso along a coastal waterway at Tortuguero on the
Caribbean coast. This 'lined-up' roosting formation is characteristic. Their pointy noses
can also be seen here. They are always near water and usually roost on tree trunks
above it. Below is a (somewhat muddy) close-up of the same species.

Carnivores aren't so easy to photograph in general, but you can get lucky. One usually fairly up-front carnivore is a coati - there are four species in the Neotropics, two of which are widespread and pretty conspicuous. They are in the same family as raccoons and some less familiar carnivores such as kinkajous. The South American Coati Nasua nasua is found throughout much of the continent and when fed can be a pest and a dangerous one; the ones at Iguazu Falls are notorious, though it's obviously not their fault. The White-nosed Coati N. narica takes its place in Central America; while I don't doubt that they could also be a bit of a problem around lodges and tourist attractions, we didn't see any evidence of it so presumably most tourists here are better educated and behaved. Unlike many carnivores they are diurnal, so we are more likely to encounter them.

Baby White-nosed Coati, part of a big group at Tapirus Lodge. These groups comprise
females and youngsters. Though I've referred to them as carnivores and they certainly hunt,
both on the ground and in trees for pretty much anything they can catch, they are
really omnivores, with fruit also being an important part of the diet.
Their erect waving tails often give them away!

Males on the other hand are solitary, except during courtship. This one was
typically on his own in the forest at Volcan Arenal.

The next one on the other hand required a lot more luck. We were returning by boat from Tortuguero (the only way to get there) and on rounding a bend came on this lovely Neotropical River Otter which had just emerged from the water with its catfish lunch. This otter has a huge range, from Mexico to Uruguay, but is threatened throughout that range, though for the most part there is insufficient knowledge about it to be sure just how much trouble it's in. It's certainly rare almost everywhere and we were very fortunate indeed. I've only seen three others, in Peru and Brazil, and they were quite fleeting and distant encounters.

Neotropical River Otter Lontra longicaudis on the Rio Suerte (ie 'Lucky River', and it was for us!)
 A rare and special viewing. They can sometimes be up to 1.5m long and weigh up to 15kg,
though that's unusual; still, it's a big animal.
And finally rodents. The most commonly encountered in Costa Rica are agoutis and squirrels, but somehow I failed to get a decent picture of the big, busy Central American Agouti Dasyprocta punctata. However to give you an idea, I'm going to inflict a decidedly below-par photo of one on you,with apologies. 
Central American Agouti foraging in the early morning at Esquinas Lodge.
They never seem to stand still! And it was this agouti which the Zoonomia project
ranked fourth in keenness of smell of the 240 mammals tested.
Agoutis are old South Americans, though not one of the originals like the sloths. Their ancestors, like the monkeys, arrived by rafting across the Atlantic from Africa. In the case of the rodents this happened some 45 million years ago (the monkeys arrived about 10 million years later).
 
Squirrels, which arrived recently from North America, weren't so difficult to find, to say the least. We saw three species, but one predominated - and if you didn't do your homework you'd be likely to think you were seeing several species! It's called the Variegated Squirrel Sciurus variegatoides for good reason. The next four photos were of this lovely big squirrel, which occurs throughout the country.
At Rincon de la Vieja, in the dry forests of the far north-west.
At Arenal Lodge, in the ranges of the central north-west - it's all-black.
 
Also at Arenal Lodge; oddly I can find no mention of the races interbreeding
but, if they're coexisting as the same species, surely they must be.

At artist Cope's garden in the central mountains.
Finally, probably my mammal highlight of the trip (though the otter was a strong competitor!). In 16 visits to South and Central America, always with skilled naturalist guides, I'd never seen a porcupine. This finally changed, within a few metres of our lunch tables at Turtle Lodge by the Caribbean at Tortuguero. It wan't easy to see to start with, but after that there was no trouble relocating it - it seemingly never moved during the three days we were there, though it must have gone foraging at night. Though their ancestors came from Africa (probably the same ancestor as the agoutis'), it wasn't the same as the African porcupines' ancestor, and they are only distantly related.
A very contented-looking Mexican Porcupine Coendou mexicanus in its favourite
bed in a palm tree. Despite its name it occurs throughout Central America.
So, that's it for a tour of some more of Costa Rica's animals. I enjoyed conducting the tour -both that one and this one - and I only hope you've enjoyed it too, otherwise there wouldn't be much point to it! Thanks for reading this far.
 
Some time I'll conclude this series with a post on the wonderful birds of Costa Rica, but next time I'll be back in Australia to introduce a lovely coastal park in my 'state of origin'. See you there.

NEXT POSTING THURSDAY 23 NOVEMBER
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